The following taxa of Dryopteris have not been treated in the preceding text. These have recently been reported by commercial growers and hobbyists, but have not been confirmed or are very recent introductions. Should the reader desire more details beyond those we have given here, we refer you to the description of the Sections and then also to appropriate floras. Most of the species from the Sino-Himalayan area were introduced into cultivation by Christopher Fraser-Jenkins and are described in Fraser-Jenkins (1989). Japanese fern species are described in English, without illustrations, in Iwatsuki et. Al. (1995). There are several recent illustrated publications on Japanese ferns, and although the text in these is in Japanese, the scientific names are usually given in Latin. We are most familiar with the eight-volume work of Karata and Nakaike that has names and indices in Latin. Volume I (1979) and Volume IV (1985) contain the Dryopteris species, hybrids inDryopteris are included in Volume VII (1994) and Volume 8 (1997) has additional distribution maps and supplementary information for taxa covered in earlier volumes.
Dryopteris blandfordii (C. Hope) C. Chr. (Index Filicum 254. 1905).Plants listed by this name were not available for verification. The name represents a large fern with fronds to ca. 90 cm tall. Mid-size plants look like D. filix-mas but are more lobed on proximal pinnules, wheareas larger plants may look like D. stewartii, but with darker stipe scales. A native of western Himalaya, Tibet, and China, the species is in Section Remotae.
Dryopteris xboottii (Tuck.) Underw. (Native Ferns, edition 4. 117. 1893).Plants listed by this name were not available for verification. The name represents a hybrid between D. intermedia (section Lophidium) and D. cristata (section Pandae). The hybrid is native to the northeastern U.S.
Dryopteris buschiana Fomin (Flora Sibiriae et Orientalis Extremi 5: 52. 1930).Plants listed under this name were not available for study and the name is of uncertain application.
"Dryopteris claytoniana," Hort.Plants listed by this name were not available for study, and valid publication of this epithet in the genus Dryopteris could not be verified. The name is probably a misspelling of Dryopteris clintoniana (which see).
"Dryopteris coreano subsp. Montana," HortPlants listed by this name were not available for study. The name is probably a misspelling for Dryopteris coreano-montana, a synonym for D. sichotensis (which see).
Dryopteris fructuosa (Christ) C. Chr. (Index Filicum 267. 1905).Plants listed by this name were not available for verification. The name represents a variable fern with markedly leathery, glossy dark green fronds to ca. 100 cm long, 2-pinnate to 2-pinnate-pinnatifid at the base. This native of the Sino-Himalayan region belongs to the section Pallidae.
Dryopteris guanchica Gibby and Jermy (Bot. J. Linn. Soc. 74: 258. 1977).Plants listed by this name were not available for verification. The name represents a plant with concolorous rhizome scales and proximal pinnae with the basal acroscopic pinnules usually shorter than the next. This native of the Canary Islands belongs to the section Lophidium.
Dryopteris koidzumiana Tagawa (Acta Phytotax. Geobot. 2: 190. 1993).This name refers to an evergreen fern similar to D. erythrosora with few scales, narrower pinnules with dentate margins, incurved teeth, and larger sori. The attractive red new fronds last longer than in D. erythrosora. Plants were introduced from Yakushima, Japan, the senior author and spores were distributed to growers. This native of Japan belongs to section Erythrovariae.
"Dryopteris megalodus," Hort.Plants listed by this name were not available for study and the name,which could not be verified as validly published, is of uncertain application.
Dryopteris munchii A. R. Smith (Proc. Calif. Acad. ser. 4, 40: 218. 1975).Plants listed by this name were not available for verification. The name represents a fern with triangular fronds to ca. 100 cm long, nearly 3-pinnate; the stipe and rhizome scales are tan with dark heavy streaks. It is native to Chiapas, Mexico and belongs to the section Cinnamomeae Fraser-Jenk. (not treated or keyed above),which is characterized by having fronds 1- to nearly 3-pinnate, lanceolate to narrowly triangular, thinly herbaceous, brittle, and the pinnules markedly obliquely sloping.
Dryopteris namegatae (Sa. Kurata) Sa. Kurata (J. Geobot. 17: 89. 1969).Plants listed by this name were not available for verification. The name represents a fern native to Japan and China that may be a hybrid between D. cycadina (D. atrata) and D. dickinsii (see treatment of D. kuratae), both belonging to section Hirtipedes.
Dryopteris odontoloma (Bedd.) C. Chr. (Acta Horti Gothob. 1: 59. 1924).Plants listed by this name were not available for verification. The name represents a fern with fronds to ca. 65 cm long, elongate triangular-lanceolate, 2-pinnate with pinnae markedly cordate at their base and small submarginal sori (to 1 mm diameter). The species is in section Pallidae.
Dryopteris pacifica (Nakai)Tagawa (Coloured Illustrations of the Japanese Pteridophyta.: 100, 211; plate 36, figure. 204. 1959).Dryopteris varia (L.)Kuntze var. hikonensis (H. Ito) Sa. Kurata.This species is currently being grown from spores by commercial and amateur growers. It is an evergreen, medium-size fern with dark green, glossy, triangular fronds to 3-pinnate and is classified in section Variae. The stipe scales are usually all blackish brown or black, the segment margins are flat, and the pinnae generally do not overlap. The indusium margin is sometimes ciliate. It is similar to D. bissetiana, but the latter has reflexed segment margins.
Dryopteris pacifica maybe confused with D. sacrosancta, but the latter has stipe scales with pale margins, lighter green fronds that are dull or hardly shiny, pinnae that tend to overlap, and indusia with entire margins. Dryopteris pacifica is evergreen and reportedly hardy in USDA zone 6 (average annual minimum temperature 0 to -10ûF). It tends to be a more compact growing plant than D. sacrosancta. Some of the plants from which spores have been distributed in U.S. were collected in Yakushima, Japan, by the senior author. The species is native to Japan, Korea and China. In Japan it grows in areas with a minimum January average temperature of 30°F or warmer.
Dryopteris paleacea (T. Moore) Hand.-Mazz. (Verh. K. K. Zool.-Bot. Ges. Wien 58:100. 1908).Plants listed by this name were not available for verification. The name is also a synonym for D. wallichiana (section Fibrillosae) and plants in the trade may represent that species (which see).
Dryopteris sordidipes Tagawa (Acta Phytotax. Geobot. 3: 29. 1934).This is an evergreen ca. 50-90 cm tall and reminiscent of D. dilatata except firmer and more coarsely cut. About 20 years ago, the name appeared in a Chicago catalog. This listing was not verified. The senior author made a recent introduction from Yakushima, Japan, and spores have been distributed to growers. It is native to Japan and Taiwan and belongs to the section Variae.
Dryopteris yigongensis Ching in C.Y. Wu (Fl. Xizangica 1: 253. 1983).Plants listed by this name were not available for verification. The name represents a fern with glossy fronds to ca. 50 cm long, narrow triangular-lanceolate, to 2-pinnate at the base and with long stipes bearing black, glossy scales. A native from the Sino-Himalayan area, it belongs to the section Fibrillosae.
Dryopteris villarii (Bell.) Woyn. ex Schinz & Thell. (Vierteljahrsschr. Naturf. Ges. Zürich 60: 339. 1915).Plants listed by this name were not available for verification. Cultivated materials may represent D. mindshelkensis (which see), a tetraploid that has D. villarii as one putative parent. It belongs to the section Pallidae.
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